Possible Clues:

• Joined the coyote chorus
• Tree Edward cut down yielded bark
• Greeted the moon
• Opening barmaid's case, I agree, was a howler
• Let out a long howl
• Made sound like hunting hounds
• Made sounds like hounds
• Ululated
• (Of a dog) howled loudly
• Made a lot of noise, but always asleep?