Related Clues:

• Howled with laughter
• Laughed loudly
• Thundered
• Bellowed
• Sounded like a lion
• Expressed approval, perhaps
• More than tittered
• Showed approval, or disapproval
• Had a big laugh
• Emulated a lion
• Shouted
• Warned, lion-style
• Split the eardrums
• Bellowed, thundered
• Made long loud sound like lion