Related Clues:

• Signified
• Relieved
• Spoke literally?
• Didn't just pronounce
• Gave a rest
• Took part in a bee
• Said 'O-D-O-U-R,' e.g.
• Bread choice
• Wheat variety
• Was in a bee, once
• Practiced with a primer
• European wheat
• Was busy at a bee?
• Joined a bee
• Like L-O-N-D-O-N