Related Clues:

• First Greek vowel
• Sharp inspiration
• Lead male or female
• Startled inhalation
• With 67 Down, pack leaders
• It's farthest from omega
• Dominant, as a male wolf
• [Heavens!]
• __ dog
• ("I'm shocked!")
• Quick intake of breath
• ___, beta, gamma
• Reaction of astonishment
• Astonished reaction
• Scared sound