Related Clues:

• Humble and obedient
• Easily alarmed, shy
• Diffident, faint-hearted
• Abjectly fearful
• After a century it's black but not brave
• Being so weak, the South appears to have been deforested
• Lacking backbone, the South seems to have become quite deforested
• Can the South be lacking trees as well as bones?
• After a century the blackbird is cowardly
• This invertebrate might end without a tree
• Me not bold blackbird comes after a century
• The South has nothing in the way of forestry, and nothing back there, either
• Being invertebrate, cut off the top to have it deforested
• There's nothing brave in having the South deforested like this
• The south could be without something like 15 across strong enough to back it