Related Clues:
- Express pain
- Fellow keeping oxen at first for beef
- Zombie sound
- Ghostly noise
- Duck caught by male grouse
- Indicate pain, discomfort, or displeasure
- Initially missed out, angry newbies complain
- Pained whine
- Rhyming partner of 'groan'
- Reacted to a joke told by 55-Across, probably
- Sound of suffering ... or pleasure
- Express unhappiness
- [Ugh, this is horrible!]
- Ghostly sound effect
- Sound in a haunted house