Possible Clues:

• Anatomical source of ill humor?
• Organ once thought to be the seat of anger
• Thus might the crescendo be placed for the height of light in days gone by
• Bad temper from stomach organ
• Body organ or bad temper
• One supplanted by general in revolution shows bad temper
• Organ associated with bad temper
• Observed to accept introduction to Paradise Lost with ill humour
• Peevishness leaving beginners one helps up
• Ductless organ